About the Authors

Chung-Yin Lee

Affiliation Histopathology Laboratory, Cancer Research UK, London Research Institute, London, United Kingdom

Rosemary Jeffery

Affiliation Histopathology Laboratory, Cancer Research UK, London Research Institute, London, United Kingdom

Gillian Hutchinson

Affiliation Biological Resources Unit, Cancer Research UK, London Research Institute, London, United Kingdom

Malcolm R. Alison

Affiliation Centre for Digestive Diseases, Blizard Institute, Barts and The London School of Medicine and Dentistry, London, United Kingdom

Richard Poulsom

Affiliations Histopathology Laboratory, Cancer Research UK, London Research Institute, London, United Kingdom, Centre for Digestive Diseases, Blizard Institute, Barts and The London School of Medicine and Dentistry, London, United Kingdom

Nicholas A. Wright

Affiliations Histopathology Laboratory, Cancer Research UK, London Research Institute, London, United Kingdom, Centre for Digestive Diseases, Blizard Institute, Barts and The London School of Medicine and Dentistry, London, United Kingdom

William R. Otto

w.r.otto@qmul.ac.uk

Affiliations Histopathology Laboratory, Cancer Research UK, London Research Institute, London, United Kingdom, Centre for Digestive Diseases, Blizard Institute, Barts and The London School of Medicine and Dentistry, London, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: WRO NAW C-YL. Performed the experiments: C-YL GH RJ RP WRO. Analyzed the data: C-YL RJ RP WRO NAW MRA. Contributed reagents/materials/analysis tools: RJ RP. Wrote the paper: WRO RP C-YL NAW MRA.