About the Authors

Naoki Yahata

Affiliations Center for iPS Cell Research and Application, Kyoto University, Kyoto, Japan, Core Research for Evolutional Science and Technology, Japan Science and Technology Agency, Saitama, Japan

Masashi Asai

Affiliations Core Research for Evolutional Science and Technology, Japan Science and Technology Agency, Saitama, Japan, Department of Pharmacology, Faculty of Medicine, Saitama Medical University, Saitama, Japan, Laboratory for Proteolytic Neuroscience, RIKEN Brain Science Institute, Saitama, Japan

Shiho Kitaoka

Affiliations Center for iPS Cell Research and Application, Kyoto University, Kyoto, Japan, Core Research for Evolutional Science and Technology, Japan Science and Technology Agency, Saitama, Japan

Kazutoshi Takahashi

Affiliation Center for iPS Cell Research and Application, Kyoto University, Kyoto, Japan

Isao Asaka

Affiliations Center for iPS Cell Research and Application, Kyoto University, Kyoto, Japan, Core Research for Evolutional Science and Technology, Japan Science and Technology Agency, Saitama, Japan

Hiroyuki Hioki

Affiliations Core Research for Evolutional Science and Technology, Japan Science and Technology Agency, Saitama, Japan, Department of Morphological Brain Science, Graduate School of Medicine, Kyoto University, Kyoto, Japan

Takeshi Kaneko

Affiliation Department of Morphological Brain Science, Graduate School of Medicine, Kyoto University, Kyoto, Japan

Kei Maruyama

Affiliation Department of Pharmacology, Faculty of Medicine, Saitama Medical University, Saitama, Japan

Takaomi C. Saido

Affiliation Laboratory for Proteolytic Neuroscience, RIKEN Brain Science Institute, Saitama, Japan

Tatsutoshi Nakahata

Affiliation Center for iPS Cell Research and Application, Kyoto University, Kyoto, Japan

Takashi Asada

Affiliation Department of Neuropsychiatry, Institute of Clinical Medicine, University of Tsukuba, Tsukuba, Japan

Shinya Yamanaka

Affiliations Center for iPS Cell Research and Application, Kyoto University, Kyoto, Japan, Yamanaka iPS Cell Special Project, Japan Science and Technology Agency, Saitama, Japan

Nobuhisa Iwata

haruhisa@cira.kyoto-u.ac.jp (HI); iwata-n@nagasaki-u.ac.jp (NI)

Affiliations Core Research for Evolutional Science and Technology, Japan Science and Technology Agency, Saitama, Japan, Laboratory for Proteolytic Neuroscience, RIKEN Brain Science Institute, Saitama, Japan, Graduate School of Biomedical Sciences, Nagasaki University, Nagasaki, Japan

Haruhisa Inoue

haruhisa@cira.kyoto-u.ac.jp (HI); iwata-n@nagasaki-u.ac.jp (NI)

Affiliations Center for iPS Cell Research and Application, Kyoto University, Kyoto, Japan, Core Research for Evolutional Science and Technology, Japan Science and Technology Agency, Saitama, Japan, Yamanaka iPS Cell Special Project, Japan Science and Technology Agency, Saitama, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: NI HI NY MA. Performed the experiments: NY MA NI. Analyzed the data: NY MA NI HI. Contributed reagents/materials/analysis tools: SK KT IA HH TK KM TCS TN TA SY. Wrote the paper: NY MA SY NI HI.