About the Authors

Gyungah Jun

Affiliations Medicine (Biomedical Genetics), Boston University Schools of Medicine and Public Health, Boston, Massachusetts, United States of America, Ophthalmology, Boston University Schools of Medicine and Public Health, Boston, Massachusetts, United States of America, Biostatistics, Boston University Schools of Medicine and Public Health, Boston, Massachusetts United States of America

Michael Nicolaou

Affiliation Medicine (Biomedical Genetics), Boston University Schools of Medicine and Public Health, Boston, Massachusetts, United States of America

Margaux A. Morrison

Affiliations Ophthalmology, Massachusetts Eye and Ear Infirmary, Boston, Massachusetts, United States of America, Ophthalmology and Visual Sciences, John A. Moran Eye Center, Center for Translational Medicine, University of Utah, Salt Lake City, Utah, United States of America

Jacqueline Buros

Affiliation Medicine (Biomedical Genetics), Boston University Schools of Medicine and Public Health, Boston, Massachusetts, United States of America

Denise J. Morgan

Affiliation Ophthalmology and Visual Sciences, John A. Moran Eye Center, Center for Translational Medicine, University of Utah, Salt Lake City, Utah, United States of America

Monte J. Radeke

Affiliation Center for the Study of Macular Degeneration, Neuroscience Research Institute, University of California Santa Barbara, Santa Barbara, California, United States of America

Yoshihiro Yonekawa

Affiliations Ophthalmology, Massachusetts Eye and Ear Infirmary, Boston, Massachusetts, United States of America, Ophthalmology, Weill Cornell Medical College, New York, New York, United States of America

Evangelia E. Tsironi

Affiliation Medicine, University of Thessaly, Larissa, Greece

Maria G. Kotoula

Affiliation Medicine, University of Thessaly, Larissa, Greece

Fani Zacharaki

Affiliation Medicine, University of Thessaly, Larissa, Greece

Nissa Mollema

Affiliation Genetics, Cell Biology, and Anatomy, University of Nebraska Medical Center, Omaha, Nebraska, United States of America

Yang Yuan

Affiliation Genetics, Cell Biology, and Anatomy, University of Nebraska Medical Center, Omaha, Nebraska, United States of America

Joan W. Miller

Affiliation Ophthalmology, Massachusetts Eye and Ear Infirmary, Boston, Massachusetts, United States of America

Neena B. Haider

Affiliation Genetics, Cell Biology, and Anatomy, University of Nebraska Medical Center, Omaha, Nebraska, United States of America

Gregory S. Hageman

Affiliation Ophthalmology and Visual Sciences, John A. Moran Eye Center, Center for Translational Medicine, University of Utah, Salt Lake City, Utah, United States of America

Ivana K. Kim

Affiliation Ophthalmology, Massachusetts Eye and Ear Infirmary, Boston, Massachusetts, United States of America

Debra A. Schaumberg

Affiliation Division of Preventive Medicine, Brigham and Women’s Hospital, Harvard Medical School, Boston, Massachusetts, United States of America

Lindsay A. Farrer

Affiliations Medicine (Biomedical Genetics), Boston University Schools of Medicine and Public Health, Boston, Massachusetts, United States of America, Ophthalmology, Boston University Schools of Medicine and Public Health, Boston, Massachusetts, United States of America, Biostatistics, Boston University Schools of Medicine and Public Health, Boston, Massachusetts United States of America, Neurology Boston University Schools of Medicine and Public Health, Boston, Massachusetts, United States of America, Epidemiology, Boston University Schools of Medicine and Public Health, Boston, Massachusetts, United States of America

Margaret M. DeAngelis

Margaret.DeAngelis@utah.edu

Affiliations Ophthalmology, Massachusetts Eye and Ear Infirmary, Boston, Massachusetts, United States of America, Ophthalmology and Visual Sciences, John A. Moran Eye Center, Center for Translational Medicine, University of Utah, Salt Lake City, Utah, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: MMD LAF GJ MJR GSH NBH. Performed the experiments: DJM MJR Y. Yonekawa FZ NM Y. Yuan. Analyzed the data: GJ MN MAM JB MJR LAF. Contributed reagents/materials/analysis tools: EET MGK JWM IKK GSH MMD LAF DAS NBH. Wrote the paper: GJ MAM MMD LAF.