About the Authors

Esther Singer

Affiliation Geomicrobiology Group, Department of Earth Sciences, University of Southern California, Los Angeles, California, United States of America

David Emerson

Affiliation Bigelow Laboratory for Ocean Sciences, West Boothbay Harbor, Maine, United States of America

Eric A. Webb

Affiliation Department of Biological Sciences, Marine Environmental Biology Section, University of Southern California, Los Angeles, California, United States of America

Roman A. Barco

Affiliation Department of Biological Sciences, Marine Environmental Biology Section, University of Southern California, Los Angeles, California, United States of America

J. Gijs Kuenen

Affiliation Department of Biotechnology, Delft University of Technology, Delft, The Netherlands

William C. Nelson

Affiliation Department of Biological Sciences, Marine Environmental Biology Section, University of Southern California, Los Angeles, California, United States of America

Clara S. Chan

Affiliation Department of Marine Chemistry and Geochemistry, Woods Hole Oceanographic Institution, Woods Hole, Massachusetts, United States of America

Luis R. Comolli

Affiliation Life Sciences Division, Lawrence Berkeley National Laboratory, Berkeley, California, United States of America

Steve Ferriera

Affiliation J. Craig Venter Institute, San Diego, California, United States of America

Justin Johnson

Affiliation J. Craig Venter Institute, San Diego, California, United States of America

John F. Heidelberg

Affiliation Department of Biological Sciences, Marine Environmental Biology Section, University of Southern California, Los Angeles, California, United States of America

Katrina J. Edwards

kje@usc.edu

Affiliations Geomicrobiology Group, Department of Earth Sciences, University of Southern California, Los Angeles, California, United States of America, Department of Biological Sciences, Marine Environmental Biology Section, University of Southern California, Los Angeles, California, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: ES DE RAB CSC LRC SF JJ. Performed the experiments: ES DE RAB CSC LRC SF JJ. Analyzed the data: ES DE EAW RAB JGK WCN CSC LRC SF JJ JFH KJE. Contributed reagents/materials/analysis tools: WCN JFH KJE. Wrote the paper: ES DE EAW JFH KJE.