About the Authors

John P. Hammond

j.hammond@nottingham.ac.uk

Affiliation Division of Plant and Crop Sciences, University of Nottingham, Loughborough, United Kingdom

Martin R. Broadley

Affiliation Division of Plant and Crop Sciences, University of Nottingham, Loughborough, United Kingdom

Helen C. Bowen

Affiliation Warwick HRI, University of Warwick, Wellesbourne, Warwick, United Kingdom

William P. Spracklen

Affiliation Warwick HRI, University of Warwick, Wellesbourne, Warwick, United Kingdom

Rory M. Hayden

Affiliation Warwick HRI, University of Warwick, Wellesbourne, Warwick, United Kingdom

Philip J. White

Affiliation The James Hutton Institute, Invergowrie, Dundee, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JPH MRB PJW. Performed the experiments: JPH HCB RMH WPS. Analyzed the data: JPH PJW MRB. Wrote the paper: JPH PJW MRB.