About the Authors

Cui Hao

Contributed equally to this work with: Cui Hao, Jiejie Hao

Affiliations Key Laboratory of Marine Drugs, Chinese Ministry of Education, Ocean University of China, Qingdao, People's Republic of China, Shandong Provincial Key Laboratory of Glycoscience and Glycotechnology, Ocean University of China, Qingdao, People's Republic of China

Jiejie Hao

Contributed equally to this work with: Cui Hao, Jiejie Hao

Affiliations Key Laboratory of Marine Drugs, Chinese Ministry of Education, Ocean University of China, Qingdao, People's Republic of China, Shandong Provincial Key Laboratory of Glycoscience and Glycotechnology, Ocean University of China, Qingdao, People's Republic of China

Wei Wang

Affiliations Key Laboratory of Marine Drugs, Chinese Ministry of Education, Ocean University of China, Qingdao, People's Republic of China, Shandong Provincial Key Laboratory of Glycoscience and Glycotechnology, Ocean University of China, Qingdao, People's Republic of China

Zhangrun Han

Affiliations Key Laboratory of Marine Drugs, Chinese Ministry of Education, Ocean University of China, Qingdao, People's Republic of China, Shandong Provincial Key Laboratory of Glycoscience and Glycotechnology, Ocean University of China, Qingdao, People's Republic of China

Guangsheng Li

Affiliations Key Laboratory of Marine Drugs, Chinese Ministry of Education, Ocean University of China, Qingdao, People's Republic of China, Shandong Provincial Key Laboratory of Glycoscience and Glycotechnology, Ocean University of China, Qingdao, People's Republic of China

Lijuan Zhang

Affiliations Key Laboratory of Marine Drugs, Chinese Ministry of Education, Ocean University of China, Qingdao, People's Republic of China, Shandong Provincial Key Laboratory of Glycoscience and Glycotechnology, Ocean University of China, Qingdao, People's Republic of China

Xia Zhao

Affiliations Key Laboratory of Marine Drugs, Chinese Ministry of Education, Ocean University of China, Qingdao, People's Republic of China, Shandong Provincial Key Laboratory of Glycoscience and Glycotechnology, Ocean University of China, Qingdao, People's Republic of China

Guangli Yu

glyu@ouc.edu.cn

Affiliations Key Laboratory of Marine Drugs, Chinese Ministry of Education, Ocean University of China, Qingdao, People's Republic of China, Shandong Provincial Key Laboratory of Glycoscience and Glycotechnology, Ocean University of China, Qingdao, People's Republic of China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: GY JH. Performed the experiments: CH JH ZH GY XZ GL. Analyzed the data: CH JH WW LZ GY. Contributed reagents/materials/analysis tools: GY JH XZ. Wrote the paper: CH WW GY LZ.