About the Authors

Lindsay E. Zanno

lzanno@fieldmuseum.org

Affiliations Field Museum of Natural History, Chicago, Illinois, United States of America, Biological Sciences Department, University of Wisconsin-Parkside, Kenosha, Wisconsin, United States of America

David J. Varricchio

Affiliation Department of Earth Sciences, Montana State University, Bozeman, Montana, United States of America

Patrick M. O'Connor

Affiliations Department of Biomedical Sciences, Ohio University College of Osteopathic Medicine, Athens, Ohio, United States of America, Ohio Center for Ecology and Evolutionary Studies, Ohio University, Athens, Ohio, United States of America

Alan L. Titus

Affiliation Grand Staircase-Escalante National Monument, Bureau of Land Management, Kanab, Utah, United States of America

Michael J. Knell

Affiliation Department of Earth Sciences, Montana State University, Bozeman, Montana, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: LEZ PMO ALT. Performed the experiments: LEZ DJV PMO ALT. Analyzed the data: LEZ DJV PMO ALT. Contributed reagents/materials/analysis tools: LEZ DJV PMO ALT MJK. Wrote the paper: LEZ DJV PMO ALT.