About the Authors

Paul R. McAdam

Affiliation The Roslin Institute and Centre for Infectious Diseases, Royal (Dick) School of Veterinary Studies, University of Edinburgh, Easter Bush, Midlothian, United Kingdom

Anne Holmes

Affiliations The Roslin Institute and Centre for Infectious Diseases, Royal (Dick) School of Veterinary Studies, University of Edinburgh, Easter Bush, Midlothian, United Kingdom, Microbiology, Royal Infirmary of Edinburgh, Edinburgh, United Kingdom

Kate E. Templeton

Affiliation Microbiology, Royal Infirmary of Edinburgh, Edinburgh, United Kingdom

J. Ross Fitzgerald

Ross.Fitzgerald@ed.ac.uk

Affiliation The Roslin Institute and Centre for Infectious Diseases, Royal (Dick) School of Veterinary Studies, University of Edinburgh, Easter Bush, Midlothian, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JRF PRM KET. Performed the experiments: PRM. Analyzed the data: JRF PRM AH KET. Wrote the paper: JRF PRM.