About the Authors

Lee A. Borthwick

Affiliation Institute of Cellular Medicine, Newcastle University, Newcastle upon Tyne, United Kingdom

Phil Botha

Affiliation Institute of Cellular Medicine, Newcastle University, Newcastle upon Tyne, United Kingdom

Bernard Verdon

Affiliation Institute for Cell & Molecular Biosciences, Newcastle University, Newcastle upon Tyne, United Kingdom

Malcolm J. Brodlie

Affiliation Institute of Cellular Medicine, Newcastle University, Newcastle upon Tyne, United Kingdom

Aaron Gardner

Affiliation Institute of Cellular Medicine, Newcastle University, Newcastle upon Tyne, United Kingdom

David Bourn

Affiliation Institute of Human Genetics, Newcastle University, Newcastle upon Tyne, United Kingdom

Gail E. Johnson

Affiliation Institute of Cellular Medicine, Newcastle University, Newcastle upon Tyne, United Kingdom

Mike A. Gray

Contributed equally to this work with: Mike A. Gray, Andrew J. Fisher

Affiliation Institute for Cell & Molecular Biosciences, Newcastle University, Newcastle upon Tyne, United Kingdom

Andrew J. Fisher

Contributed equally to this work with: Mike A. Gray, Andrew J. Fisher

A.J.Fisher@newcastle.ac.uk

Affiliation Institute of Cellular Medicine, Newcastle University, Newcastle upon Tyne, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: LAB PB MAG AJF. Performed the experiments: LAB PB BV DB GEJ. Analyzed the data: LAB PB BV AG. Contributed reagents/materials/analysis tools: BV MJB. Wrote the paper: LAB MAG AJF. Figure preparation: AG.