About the Authors

Naoko Iguchi

Affiliation Monell Chemical Senses Center, Philadelphia, Pennsylvania, United States of America

Tadahiro Ohkuri

Affiliation Monell Chemical Senses Center, Philadelphia, Pennsylvania, United States of America

Jay P. Slack

Affiliation Givaudan Flavors Corporation, Cincinnati, Ohio, United States of America

Ping Zhong

Affiliation Givaudan Flavors Corporation, Cincinnati, Ohio, United States of America

Liquan Huang

lhuang@monell.org

Affiliation Monell Chemical Senses Center, Philadelphia, Pennsylvania, United States of America

Competing Interests

NI and LH have declared that no competing interests exist. JPS and PZ are employees of Givaudan Flavors Corp. This does not alter the authors' adherence to all the PLoS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: LH PZ NI JPS TO. Performed the experiments: NI TO JPS. Analyzed the data: LH NI TO JPS PZ. Contributed reagents/materials/analysis tools: LH NI TO JPS. Wrote the paper: NI LH TO PZ JPS.