About the Authors

Sanae Sakai

sakai-s@jamstec.go.jp

Affiliations Institute of Biogeosciences, Japan Agency for Marine-Earth Science and Technology (JAMSTEC), Yokosuka, Kanagawa, Japan, Department of Environmental Systems Engineering, Nagaoka University of Technology, Nagaoka, Niigata, Japan

Yoshihiro Takaki

Affiliation Institute of Biogeosciences, Japan Agency for Marine-Earth Science and Technology (JAMSTEC), Yokosuka, Kanagawa, Japan

Shigeru Shimamura

Affiliation Institute of Biogeosciences, Japan Agency for Marine-Earth Science and Technology (JAMSTEC), Yokosuka, Kanagawa, Japan

Mitsuo Sekine

Affiliation Biological Resource Center, National Institute of Technology and Evaluation (NITE), Shibuya-ku, Tokyo, Japan

Takahisa Tajima

Affiliation Biological Resource Center, National Institute of Technology and Evaluation (NITE), Shibuya-ku, Tokyo, Japan

Hiroki Kosugi

Affiliation Biological Resource Center, National Institute of Technology and Evaluation (NITE), Shibuya-ku, Tokyo, Japan

Natsuko Ichikawa

Affiliation Biological Resource Center, National Institute of Technology and Evaluation (NITE), Shibuya-ku, Tokyo, Japan

Eiji Tasumi

Affiliation Biological Resource Center, National Institute of Technology and Evaluation (NITE), Shibuya-ku, Tokyo, Japan

Aiko T. Hiraki

Affiliation Biological Resource Center, National Institute of Technology and Evaluation (NITE), Shibuya-ku, Tokyo, Japan

Ai Shimizu

Affiliation Biological Resource Center, National Institute of Technology and Evaluation (NITE), Shibuya-ku, Tokyo, Japan

Yumiko Kato

Affiliation Biological Resource Center, National Institute of Technology and Evaluation (NITE), Shibuya-ku, Tokyo, Japan

Rika Nishiko

Affiliation Biological Resource Center, National Institute of Technology and Evaluation (NITE), Shibuya-ku, Tokyo, Japan

Koji Mori

Affiliation Biological Resource Center, National Institute of Technology and Evaluation (NITE), Shibuya-ku, Tokyo, Japan

Nobuyuki Fujita

Affiliation Biological Resource Center, National Institute of Technology and Evaluation (NITE), Shibuya-ku, Tokyo, Japan

Hiroyuki Imachi

Affiliation Institute of Biogeosciences, Japan Agency for Marine-Earth Science and Technology (JAMSTEC), Yokosuka, Kanagawa, Japan

Ken Takai

Affiliation Institute of Biogeosciences, Japan Agency for Marine-Earth Science and Technology (JAMSTEC), Yokosuka, Kanagawa, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: S. Sakai YT HI KT. Performed the experiments: S. Sakai MS HK YK RN. Analyzed the data: YT S. Sakai TT NI ET ATH AS KM. Contributed reagents/materials/analysis tools: YT S. Shimamura. Wrote the paper: S. Sakai YT HI KT. Coordination of the project: HI NF.