About the Authors

Michèle M. Sale

msale@virginia.edu

Affiliations Center for Public Health Genomics, University of Virginia, Charlottesville, Virginia, United States of America, Department of Medicine, University of Virginia, Charlottesville, Virginia, United States of America, Department of Biochemistry and Molecular Genetics, University of Virginia, Charlottesville, Virginia, United States of America

Wei-Min Chen

Affiliations Center for Public Health Genomics, University of Virginia, Charlottesville, Virginia, United States of America, Department of Public Health Sciences, University of Virginia, Charlottesville, Virginia, United States of America

Daniel E. Weeks

Affiliations Department of Human Genetics, Graduate School of Public Health, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America, Department of Biostatistics, Graduate School of Public Health, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

Josyf C. Mychaleckyj

Affiliations Center for Public Health Genomics, University of Virginia, Charlottesville, Virginia, United States of America, Department of Public Health Sciences, University of Virginia, Charlottesville, Virginia, United States of America

Xuanlin Hou

Affiliations Center for Public Health Genomics, University of Virginia, Charlottesville, Virginia, United States of America, Department of Public Health Sciences, University of Virginia, Charlottesville, Virginia, United States of America

Miranda Marion

Affiliation Department of Biostatistical Sciences, Wake Forest University School of Medicine, Winston-Salem, North Carolina, United States of America

Fernando Segade

Affiliation Department of Anatomy and Cell Biology, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Margaretha L. Casselbrant

Affiliations Department of Human Genetics, Graduate School of Public Health, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America, Department of Otolaryngology, Children's Hospital of Pittsburgh of UPMC, Pittsburgh, Pennsylvania, United States of America

Ellen M. Mandel

Affiliation Department of Otolaryngology, Children's Hospital of Pittsburgh of UPMC, Pittsburgh, Pennsylvania, United States of America

Robert E. Ferrell

Affiliation Department of Human Genetics, Graduate School of Public Health, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

Stephen S. Rich

Affiliations Center for Public Health Genomics, University of Virginia, Charlottesville, Virginia, United States of America, Department of Medicine, University of Virginia, Charlottesville, Virginia, United States of America, Department of Public Health Sciences, University of Virginia, Charlottesville, Virginia, United States of America

Kathleen A. Daly

Affiliation Department of Otolaryngology, University of Minnesota, Minneapolis, Minnesota, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: MMS FS SSR KAD. Performed the experiments: MMS REF. Analyzed the data: W-MC XH DEW MM. Contributed reagents/materials/analysis tools: JCM XH MLC EMM KAD. Wrote the paper: MMS W-MC DEW SSR KAD.