About the Authors

Chen-Yi Wang

Affiliation Institute of Cellular and Organismic Biology, Academia Sinica, Taipei, Taiwan

Chia-Yin Hua

Contributed equally to this work with: Chia-Yin Hua, Hsiang-En Hsu

Affiliation Institute of Cellular and Organismic Biology, Academia Sinica, Taipei, Taiwan

Hsiang-En Hsu

Contributed equally to this work with: Chia-Yin Hua, Hsiang-En Hsu

Affiliation Institute of Cellular and Organismic Biology, Academia Sinica, Taipei, Taiwan

Chia-Ling Hsu

Affiliation Institute of Cellular and Organismic Biology, Academia Sinica, Taipei, Taiwan

Hsin-Yi Tseng

Affiliation Institute of Cellular and Organismic Biology, Academia Sinica, Taipei, Taiwan

Duncan E. Wright

Affiliation Institute of Cellular and Organismic Biology, Academia Sinica, Taipei, Taiwan

Pang-Hung Hsu

Affiliations Institute of Biological Chemistry, Academia Sinica, Taipei, Taiwan, Department of Life Science, Institute of Bioscience and Biotechnology, National Taiwan Ocean University, Keelung, Taiwan

Chih-Hung Jen

Current address: Xinscience Biotech Co., Ltd, Taipei City, Taiwan

Affiliation VGH Yang-Ming Genome Research Center, Taipei, Taiwan

Chia-Yeh Lin

Affiliation Institute of Cellular and Organismic Biology, Academia Sinica, Taipei, Taiwan

Meng-Ying Wu

Affiliation Institute of Cellular and Organismic Biology, Academia Sinica, Taipei, Taiwan

Min-Daw Tsai

Affiliation Institute of Biological Chemistry, Academia Sinica, Taipei, Taiwan

Cheng-Fu Kao

ckao@gate.sinica.edu.tw

Affiliation Institute of Cellular and Organismic Biology, Academia Sinica, Taipei, Taiwan

Competing Interests

CHJ was affiliated with National Yang-Ming University and subsequently moved to Xinscience Biotech. This company plays no role in the study. This does not alter the authors' adherence to all the PLoS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: CFK MDT. Performed the experiments: CYW CYH HEH HYT CLH PHH. Analyzed the data: PHH CHJ. Contributed reagents/materials/analysis tools: CYL MYW. Wrote the paper: CFK CYW CLH DEW.