About the Authors

Yi Zhong

Contributed equally to this work with: Yi Zhong, Zheng Wang

Affiliation Department of Pathology, The Sol Goldman Pancreatic Cancer Research Center, Johns Hopkins Medical Institutions, Baltimore, Maryland, United States of America

Zheng Wang

Contributed equally to this work with: Yi Zhong, Zheng Wang

Affiliations Department of Pathology, The Sol Goldman Pancreatic Cancer Research Center, Johns Hopkins Medical Institutions, Baltimore, Maryland, United States of America, Department of Hepatobiliary Surgery, The First Affiliated Hospital, School of Medicine of Xi'an Jiaotong University, Shanxi, China

Baojin Fu

Affiliation Department of Pathology, The Sol Goldman Pancreatic Cancer Research Center, Johns Hopkins Medical Institutions, Baltimore, Maryland, United States of America

Fan Pan

Affiliation Department of Oncology, The Sol Goldman Pancreatic Cancer Research Center, Johns Hopkins Medical Institutions, Baltimore, Maryland, United States of America

Shinichi Yachida

Affiliation Department of Pathology, The Sol Goldman Pancreatic Cancer Research Center, Johns Hopkins Medical Institutions, Baltimore, Maryland, United States of America

Mousumi Dhara

Affiliation Department of Pathology, The Sol Goldman Pancreatic Cancer Research Center, Johns Hopkins Medical Institutions, Baltimore, Maryland, United States of America

Emilia Albesiano

Affiliation Department of Oncology, The Sol Goldman Pancreatic Cancer Research Center, Johns Hopkins Medical Institutions, Baltimore, Maryland, United States of America

Li Li

Affiliations Department of Pathology, The Sol Goldman Pancreatic Cancer Research Center, Johns Hopkins Medical Institutions, Baltimore, Maryland, United States of America, Department of Opthamology, The First Affiliated Hospital, School of Medicine of Xi'an Jiaotong University, Shanxi, China

Yoshiki Naito

Affiliation Department of Pathology, The Sol Goldman Pancreatic Cancer Research Center, Johns Hopkins Medical Institutions, Baltimore, Maryland, United States of America

Felip Vilardell

Affiliations Department of Pathology, The Sol Goldman Pancreatic Cancer Research Center, Johns Hopkins Medical Institutions, Baltimore, Maryland, United States of America, Hospital Universitari Arnau de Vilanova, Lleida, Spain

Christopher Cummings

Affiliations Department of Pathology, The Sol Goldman Pancreatic Cancer Research Center, Johns Hopkins Medical Institutions, Baltimore, Maryland, United States of America, University of Colorado School of Medicine, Denver, Colorado, United States of America

Paola Martinelli

Affiliation Programa de Patologia Molecular, Centro Nacional de Investigaciones Oncológicas, Madrid, Spain

Ang Li

Affiliation Department of Pathology, The Sol Goldman Pancreatic Cancer Research Center, Johns Hopkins Medical Institutions, Baltimore, Maryland, United States of America

Raluca Yonescu

Affiliation Department of Pathology, The Sol Goldman Pancreatic Cancer Research Center, Johns Hopkins Medical Institutions, Baltimore, Maryland, United States of America

Qingyong Ma

Affiliation Department of Hepatobiliary Surgery, The First Affiliated Hospital, School of Medicine of Xi'an Jiaotong University, Shanxi, China

Constance A. Griffin

Affiliations Department of Pathology, The Sol Goldman Pancreatic Cancer Research Center, Johns Hopkins Medical Institutions, Baltimore, Maryland, United States of America, Department of Oncology, The Sol Goldman Pancreatic Cancer Research Center, Johns Hopkins Medical Institutions, Baltimore, Maryland, United States of America

Francisco X. Real

Affiliations Programa de Patologia Molecular, Centro Nacional de Investigaciones Oncológicas, Madrid, Spain, Departament de Ciències Experimentals i de la Salut, Universitat Pompeu Fabra, Barcelona, Spain

Christine A. Iacobuzio-Donahue

ciacobu@jhmi.edu

Affiliations Department of Pathology, The Sol Goldman Pancreatic Cancer Research Center, Johns Hopkins Medical Institutions, Baltimore, Maryland, United States of America, Department of Oncology, The Sol Goldman Pancreatic Cancer Research Center, Johns Hopkins Medical Institutions, Baltimore, Maryland, United States of America, Department of Surgery, The Sol Goldman Pancreatic Cancer Research Center, Johns Hopkins Medical Institutions, Baltimore, Maryland, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: YZ ZW BF FP EA PM QM CAG FXR CAI-D. Performed the experiments: YZ ZW BF FP SY MD EA LL YN FV CC AL RY. Analyzed the data: YZ ZW BF FP SY MD EA LL YN FV RY CAI-D. Contributed reagents/materials/analysis tools: YZ ZW BF FP SY MD EA LL YN FV CC AL RY QM CAG FXR CAI-D. Wrote the paper: YZ ZW BF CAI-D.