About the Authors

Sanae Hamanaka

Affiliations Japan Science Technology Agency, Exploratory Research for Advanced Technology (ERATO), Nakauchi Stem Cell and Organ Regeneration Project, Tokyo, Japan, Division of Stem Cell Therapy, Center for Stem Cell Biology and Regenerative Medicine, Institute of Medical Science, University of Tokyo, Tokyo, Japan

Tomoyuki Yamaguchi

tomoyama@ims.u-tokyo.ac.jp (TY); nakauchi@ims.u-tokyo.ac.jp (HN)

Affiliations Japan Science Technology Agency, Exploratory Research for Advanced Technology (ERATO), Nakauchi Stem Cell and Organ Regeneration Project, Tokyo, Japan, Division of Stem Cell Therapy, Center for Stem Cell Biology and Regenerative Medicine, Institute of Medical Science, University of Tokyo, Tokyo, Japan

Toshihiro Kobayashi

Affiliations Japan Science Technology Agency, Exploratory Research for Advanced Technology (ERATO), Nakauchi Stem Cell and Organ Regeneration Project, Tokyo, Japan, Division of Stem Cell Therapy, Center for Stem Cell Biology and Regenerative Medicine, Institute of Medical Science, University of Tokyo, Tokyo, Japan

Megumi Kato-Itoh

Affiliation Division of Stem Cell Therapy, Center for Stem Cell Biology and Regenerative Medicine, Institute of Medical Science, University of Tokyo, Tokyo, Japan

Satoshi Yamazaki

Affiliation Division of Stem Cell Therapy, Center for Stem Cell Biology and Regenerative Medicine, Institute of Medical Science, University of Tokyo, Tokyo, Japan

Hideyuki Sato

Affiliation Division of Stem Cell Therapy, Center for Stem Cell Biology and Regenerative Medicine, Institute of Medical Science, University of Tokyo, Tokyo, Japan

Ayumi Umino

Affiliation Division of Stem Cell Therapy, Center for Stem Cell Biology and Regenerative Medicine, Institute of Medical Science, University of Tokyo, Tokyo, Japan

Yukiko Wakiyama

Affiliation Division of Stem Cell Therapy, Center for Stem Cell Biology and Regenerative Medicine, Institute of Medical Science, University of Tokyo, Tokyo, Japan

Mami Arai

Affiliation Division of Stem Cell Therapy, Center for Stem Cell Biology and Regenerative Medicine, Institute of Medical Science, University of Tokyo, Tokyo, Japan

Makoto Sanbo

Affiliation Center for Genetic Analysis of Behavior, National Institute for Physiological Sciences, Okazaki, Japan

Masumi Hirabayashi

Affiliations Center for Genetic Analysis of Behavior, National Institute for Physiological Sciences, Okazaki, Japan, School of Life Science, The Graduate University for Advanced Studies, Okazaki, Japan

Hiromitsu Nakauchi

tomoyama@ims.u-tokyo.ac.jp (TY); nakauchi@ims.u-tokyo.ac.jp (HN)

Affiliations Japan Science Technology Agency, Exploratory Research for Advanced Technology (ERATO), Nakauchi Stem Cell and Organ Regeneration Project, Tokyo, Japan, Division of Stem Cell Therapy, Center for Stem Cell Biology and Regenerative Medicine, Institute of Medical Science, University of Tokyo, Tokyo, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SH TY HN. Performed the experiments: SH TY MH MK-I MS TK AU HS YW. Analyzed the data: SH TK SY. Contributed reagents/materials/analysis tools: TY MA. Wrote the paper: SH TY. Final approval of the manuscript: TY HN.