About the Authors

Kristine Bohmann

Affiliation Centre for GeoGenetics, Natural History Museum of Denmark, Copenhagen, Denmark

Ara Monadjem

Affiliation All Out Africa Research Unit, Department of Biological Sciences, University of Swaziland, Kwaluseni, Swaziland

Christina Lehmkuhl Noer

Affiliation Centre for GeoGenetics, Natural History Museum of Denmark, Copenhagen, Denmark

Morten Rasmussen

Affiliation Centre for GeoGenetics, Natural History Museum of Denmark, Copenhagen, Denmark

Matt R. K. Zeale

Affiliation School of Biological Sciences, University of Bristol, Bristol, United Kingdom

Elizabeth Clare

Affiliation School of Biological Sciences, University of Bristol, Bristol, United Kingdom

Gareth Jones

Affiliation School of Biological Sciences, University of Bristol, Bristol, United Kingdom

Eske Willerslev

Affiliation Centre for GeoGenetics, Natural History Museum of Denmark, Copenhagen, Denmark

M. Thomas P. Gilbert

mtpgilbert@gmail.com

Affiliation Centre for GeoGenetics, Natural History Museum of Denmark, Copenhagen, Denmark

Competing Interests

Field assistance for this study was provided by the Royal Swaziland Sugar Corporation. Author Tom Gilbert is an academic editor for PLoS ONE. This does not alter the authors' adherence to all the PLoS ONE policies on sharing data and material.

Author Contributions

Conceived and designed the experiments: KB AM CLN MTPG. Performed the experiments: KB AM CLN MR MTPG. Analyzed the data: KM AM CLN MR MZ EC MTPG. Contributed reagents/materials/analysis tools: AM EW MTPG. Wrote the paper: KW AM MZ EC GJ EW MTPG.