About the Authors

Elyssa L. Barron

Affiliation Laboratory of Infectious Diseases, National Institute of Allergy and Infectious Diseases, National Institutes of Health, Department of Health and Human Services, Bethesda, Maryland, United States of America

Stanislav V. Sosnovtsev

Affiliation Laboratory of Infectious Diseases, National Institute of Allergy and Infectious Diseases, National Institutes of Health, Department of Health and Human Services, Bethesda, Maryland, United States of America

Karin Bok

Affiliation Laboratory of Infectious Diseases, National Institute of Allergy and Infectious Diseases, National Institutes of Health, Department of Health and Human Services, Bethesda, Maryland, United States of America

Victor Prikhodko

Affiliation Laboratory of Infectious Diseases, National Institute of Allergy and Infectious Diseases, National Institutes of Health, Department of Health and Human Services, Bethesda, Maryland, United States of America

Carlos Sandoval-Jaime

Affiliation Laboratory of Infectious Diseases, National Institute of Allergy and Infectious Diseases, National Institutes of Health, Department of Health and Human Services, Bethesda, Maryland, United States of America

Crystal R. Rhodes

Affiliation Laboratory of Infectious Diseases, National Institute of Allergy and Infectious Diseases, National Institutes of Health, Department of Health and Human Services, Bethesda, Maryland, United States of America

Kim Hasenkrug

Affiliation Retrovirus Immunology Section, National Institute of Allergy and Infectious Diseases, National Institutes of Health, Department of Health and Human Services, Hamilton, Montana, United States of America

Aaron B. Carmody

Affiliation Retrovirus Immunology Section, National Institute of Allergy and Infectious Diseases, National Institutes of Health, Department of Health and Human Services, Hamilton, Montana, United States of America

Jerrold M. Ward

Affiliation Comparative Medicine Branch, National Institute of Allergy and Infectious Diseases, National Institutes of Health, Department of Health and Human Services, Bethesda, Maryland, United States of America

Kathy Perdue

Affiliation Comparative Medicine Branch, National Institute of Allergy and Infectious Diseases, National Institutes of Health, Department of Health and Human Services, Bethesda, Maryland, United States of America

Kim Y. Green

kgreen@niaid.nih.gov

Affiliation Laboratory of Infectious Diseases, National Institute of Allergy and Infectious Diseases, National Institutes of Health, Department of Health and Human Services, Bethesda, Maryland, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: ELB SVS KB JMW KP KYG. Performed the experiments: ELB SVS KB VP CS-J CRR JMW KP KYG. Analyzed the data: ELB SVS KB JMW KP KYG. Contributed reagents/materials/analysis tools: KH ABC. Wrote the paper: ELB KYG. Supervision of veterinary technical staff: KP.