About the Authors

Karen Hudson

Current address: Crop Production and Pest Control Research Unit, Agricultural Research Service, United States Department of Agriculture, West Lafayette, Indiana, United States of America

Affiliation Howard Hughes Medical Institute, Department of Molecular Genetics and Cell Biology, University of Chicago, Chicago, Illinois, United States of America

Song Luo

Current address: Chromatin Inc, Chicago, Illinois, United States of America

Affiliation Howard Hughes Medical Institute, Department of Molecular Genetics and Cell Biology, University of Chicago, Chicago, Illinois, United States of America

Nicole Hagemann

Affiliation Howard Hughes Medical Institute, Department of Molecular Genetics and Cell Biology, University of Chicago, Chicago, Illinois, United States of America

Daphne Preuss

dpreuss@chromatininc.com

Current address: Chromatin Inc, Chicago, Illinois, United States of America

Affiliation Howard Hughes Medical Institute, Department of Molecular Genetics and Cell Biology, University of Chicago, Chicago, Illinois, United States of America

Competing Interests

Authors SL and DP are currently employed by Chromatin, Inc. This does not alter the authors' adherence to the PLoS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: SL DP. Performed the experiments: SL NH KH. Analyzed the data: KH. Wrote the paper: KH.