About the Authors

Fusheng Li

Affiliation Vaccine and Infectious Disease Division, Fred Hutchinson Cancer Research Center, Seattle, Washington, United States of America

Adam C. Finnefrock

Affiliation Vaccine Basic Research Department, Merck Research Laboratories, West Point, Pennsylvania, United States of America

Sheri A. Dubey

Affiliation Vaccine Basic Research Department, Merck Research Laboratories, West Point, Pennsylvania, United States of America

Bette T. M. Korber

Affiliations Santa Fe Institute, Santa Fe, New Mexico, United States of America, Los Alamos Laboratory, Los Alamos, New Mexico, United States of America

James Szinger

Affiliation Los Alamos Laboratory, Los Alamos, New Mexico, United States of America

Suzanne Cole

Affiliation Vaccine Basic Research Department, Merck Research Laboratories, West Point, Pennsylvania, United States of America

M. Juliana McElrath

Affiliations Vaccine and Infectious Disease Division, Fred Hutchinson Cancer Research Center, Seattle, Washington, United States of America, Department of Medicine, University of Washington, Seattle, Washington, United States of America

John W. Shiver

Affiliation Vaccine Basic Research Department, Merck Research Laboratories, West Point, Pennsylvania, United States of America

Danilo R. Casimiro

Affiliation Vaccine Basic Research Department, Merck Research Laboratories, West Point, Pennsylvania, United States of America

Lawrence Corey

Affiliations Vaccine and Infectious Disease Division, Fred Hutchinson Cancer Research Center, Seattle, Washington, United States of America, Department of Medicine, University of Washington, Seattle, Washington, United States of America, Department of Laboratory Medicine, University of Washington, Seattle, Washington, United States of America, Department of Epidemiology, University of Washington, Seattle, Washington, United States of America

Steven G. Self

sgs@scharp.org

Affiliations Vaccine and Infectious Disease Division, Fred Hutchinson Cancer Research Center, Seattle, Washington, United States of America, Department of Biostatistics, University of Washington, Seattle, Washington, United States of America

Competing Interests

Some authors from Merck Research Laboratories have competing interests. This does not alter the authors' adherence to all the PLoS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: ACF SAD JWS DRC. Performed the experiments: SAD SC. Analyzed the data: LF ACF BTMK JS. Wrote the paper: FL ACF SAD LC SGS MJM. Developed methods: LF ACF BTMK JS.