About the Authors

Ann L. Oberg

Affiliation Department of Health Sciences Research, Mayo Clinic, Rochester, Minnesota, United States of America

Amy J. French

Affiliation Department of Laboratory Medicine and Pathology, Mayo Clinic, Rochester, Minnesota, United States of America

Aaron L. Sarver

Affiliation Biostatistics and Informatics, Masonic Cancer Center, University of Minnesota, Minneapolis, Minnesota, United States of America

Subbaya Subramanian

Affiliation Department of Surgery, University of Minnesota, Minneapolis, Minnesota, United States of America

Bruce W. Morlan

Affiliation Department of Health Sciences Research, Mayo Clinic, Rochester, Minnesota, United States of America

Shaun M. Riska

Affiliation Department of Health Sciences Research, Mayo Clinic, Rochester, Minnesota, United States of America

Pedro M. Borralho

Affiliation iMed.UL, Faculty of Pharmacy, University of Lisbon, Lisbon, Portugal

Julie M. Cunningham

Affiliation Department of Laboratory Medicine and Pathology, Mayo Clinic, Rochester, Minnesota, United States of America

Lisa A. Boardman

Affiliation Division of Gastroenterology and Hepatology, Mayo Clinic, Rochester, Minnesota, United States of America

Liang Wang

Affiliation Department of Laboratory Medicine and Pathology, Mayo Clinic, Rochester, Minnesota, United States of America

Thomas C. Smyrk

Affiliation Department of Laboratory Medicine and Pathology, Mayo Clinic, Rochester, Minnesota, United States of America

Yan Asmann

Affiliation Department of Health Sciences Research, Mayo Clinic, Rochester, Minnesota, United States of America

Clifford J. Steer

Affiliation Department of Medicine, and Department of Genetics, Cell Biology and Development, University of Minnesota, Minneapolis, Minnesota, United States of America

Stephen N. Thibodeau

sthibodeau@mayo.edu

Affiliation Department of Laboratory Medicine and Pathology, Mayo Clinic, Rochester, Minnesota, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: ALO AJF ALS SS BM SMR PMB JMC LAB LW TCS YA CJS SNT. Performed the experiments: AJF PMB JMC TCS LW. Analyzed the data: ALO ALS BM SMR AJF SS YA. Contributed reagents/materials/analysis tools: SNT CJS. Wrote the paper: ALO AJF ALS SS CJS SNT.