About the Authors

Ting Lei

Affiliation Department of Veterinary Biomedical Sciences, College of Veterinary Medicine, University of Minnesota, St. Paul, Minnesota, United States of America

Xudong Liang

Affiliation Department of Veterinary Biomedical Sciences, College of Veterinary Medicine, University of Minnesota, St. Paul, Minnesota, United States of America

Junshu Yang

Affiliation Department of Veterinary Biomedical Sciences, College of Veterinary Medicine, University of Minnesota, St. Paul, Minnesota, United States of America

Meiying Yan

Affiliation Department of Veterinary Biomedical Sciences, College of Veterinary Medicine, University of Minnesota, St. Paul, Minnesota, United States of America

Li Zheng

Affiliation Department of Veterinary Biomedical Sciences, College of Veterinary Medicine, University of Minnesota, St. Paul, Minnesota, United States of America

Bruce Walcheck

Affiliation Department of Veterinary Biomedical Sciences, College of Veterinary Medicine, University of Minnesota, St. Paul, Minnesota, United States of America

Yinduo Ji

jixxx002@umn.edu

Affiliation Department of Veterinary Biomedical Sciences, College of Veterinary Medicine, University of Minnesota, St. Paul, Minnesota, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: TL BW YJ. Performed the experiments: TL XL JY MY LZ BW. Analyzed the data: TL BW YJ. Wrote the paper: TL YJ.