About the Authors

Tiffani A. Greene

Affiliation Integral Molecular, Inc., Philadelphia, Pennsylvania, United States of America

Suzanne Alarcon

Affiliation Department of Nutritional Sciences, School of Environmental and Biological Sciences, Rutgers University, New Brunswick, New Jersey, United States of America

Anu Thomas

Affiliation Integral Molecular, Inc., Philadelphia, Pennsylvania, United States of America

Eli Berdougo

Affiliation Integral Molecular, Inc., Philadelphia, Pennsylvania, United States of America

Benjamin J. Doranz

Affiliation Integral Molecular, Inc., Philadelphia, Pennsylvania, United States of America

Paul A. S. Breslin

Affiliations Department of Nutritional Sciences, School of Environmental and Biological Sciences, Rutgers University, New Brunswick, New Jersey, United States of America, Monell Chemical Senses Center, Philadelphia, Pennsylvania, United States of America

Joseph B. Rucker

jrucker@integralmolecular.com

Affiliation Integral Molecular, Inc., Philadelphia, Pennsylvania, United States of America

Competing Interests

P.A.S.B. and S.A. have declared that no competing interests exist. T.A.G., A.T., E.B., B.J.D., and J.B.R. are all employees of Integral Molecular. B.J.D. and J.B.R. are shareholders of Integral Molecular. Integral Molecular has filed a provisional patent on probenecid as a bitter taste inhibitor. This does not alter the authors' adherence to all the PLoS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: TAG BJD PASB JBR. Performed the experiments: TAG SA AT. Analyzed the data: TAG SA AT EB PASB JBR. Wrote the paper: EB BJD PASB JBR.