About the Authors

Daniel D. Shaye

ds451@columbia.edu (DDS); isg4@columbia.edu (IG)

Affiliation Howard Hughes Medical Institute, Columbia University, College of Physicians and Surgeons, New York, New York, United States of America

Iva Greenwald

ds451@columbia.edu (DDS); isg4@columbia.edu (IG)

Affiliations Howard Hughes Medical Institute, Columbia University, College of Physicians and Surgeons, New York, New York, United States of America, Department of Biochemistry and Molecular Biophysics, Columbia University, College of Physicians and Surgeons, New York, New York, United States of America, Department of Genetics and Development, Columbia University, College of Physicians and Surgeons, New York, New York, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: DDS IG. Performed the experiments: DDS. Analyzed the data: DDS. Contributed reagents/materials/analysis tools: DDS. Wrote the paper: DDS IG.