About the Authors

Samantha L. Bell

Affiliation Department of Biological Sciences, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

Annette N. Chiang

Affiliation Department of Biological Sciences, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

Jeffrey L. Brodsky

jbrodsky@pitt.edu

Affiliation Department of Biological Sciences, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JLB SLB ANC. Performed the experiments: SLB ANC. Analyzed the data: SLB ANC JLB. Contributed reagents/materials/analysis tools: SLB ANC. Wrote the paper: SLB JLB.