About the Authors

Vibhor Kumar

Affiliations Department of Biomedical Engineering and Computational Science, School of Science and Technology, Centre of Excellence in Computational Complex Systems Research, Aalto University Aalto, Finland, Computational and Mathematical Biology, Genome Institute of Singapore, A*STAR, Singapore

Sarah J. Butcher

Affiliation Institute of Biotechnology, University of Helsinki, Helsinki, Finland

Katariina Öörni

Affiliation Wihuri Research Institute, Kalliolinnantie 4, Helsinki, Finland

Peter Engelhardt

Affiliations Department of Biomedical Engineering and Computational Science, School of Science and Technology, Centre of Excellence in Computational Complex Systems Research, Aalto University Aalto, Finland, Department of Pathology, Haartman Institute, Haartmaninkatu 3, University of Helsinki, Helsinki, Finland, Department of Applied Physics, Nanomicroscopy Center, School of Science and Technology, Puumiehenkuja 2, Aalto University, Espoo, Finland

Jukka Heikkonen

Affiliations Department of Biomedical Engineering and Computational Science, School of Science and Technology, Centre of Excellence in Computational Complex Systems Research, Aalto University Aalto, Finland, Department of Information Technology, University of Turku, Turku, Finland

Kimmo Kaski

Affiliation Department of Biomedical Engineering and Computational Science, School of Science and Technology, Centre of Excellence in Computational Complex Systems Research, Aalto University Aalto, Finland

Mika Ala-Korpela

Affiliations Computational Medicine Research Group, Institute of Clinical Medicine, Faculty of Medicine, Biocenter Oulu, University of Oulu, Oulu, Finland, Department of Internal Medicine and Biocenter Oulu, Clinical Research Center, University of Oulu, Oulu, Finland

Petri T. Kovanen

petri.kovanen@wri.fi

Affiliation Wihuri Research Institute, Kalliolinnantie 4, Helsinki, Finland

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: VK SJB KÖ PE JH KK MA-K PTK. Performed the experiments: SJB KÖ. Analyzed the data: VK SJB PE JH. Wrote the paper: VK SJB KÖ MA-K PTK.