About the Authors

Galliano Zanello

Affiliations Société Industrielle Lesaffre, Lesaffre Feed Additives, Marcq-en-Baroeul, France, Institut National de la Recherche Agronomique (INRA), UR1282, Infectiologie Animale et Santé Publique, Nouzilly (Tours), France

Mustapha Berri

Affiliation Institut National de la Recherche Agronomique (INRA), UR1282, Infectiologie Animale et Santé Publique, Nouzilly (Tours), France

Joëlle Dupont

Affiliation Institut National de la Recherche Agronomique (INRA), UMR85, Physiologie de la Reproduction et des Comportements, Nouzilly (Tours), France

Pierre-Yves Sizaret

Affiliation Département des microscopies, plate-forme R.I.O de microscopie électronique, Université François Rabelais, Tours, France

Romain D'Inca

Affiliation Société Industrielle Lesaffre, Lesaffre Feed Additives, Marcq-en-Baroeul, France

Henri Salmon

Contributed equally to this work with: Henri Salmon, François Meurens

Affiliation Institut National de la Recherche Agronomique (INRA), UR1282, Infectiologie Animale et Santé Publique, Nouzilly (Tours), France

François Meurens

Contributed equally to this work with: Henri Salmon, François Meurens

meurens@tours.inra.fr

Affiliation Institut National de la Recherche Agronomique (INRA), UR1282, Infectiologie Animale et Santé Publique, Nouzilly (Tours), France

Competing Interests

GZ and RDI are employees of Lesaffre. Lesaffre has a patent application related to strain CNCM I-3856, which is a commercial strain. This does not alter the authors' adherence to all the PLoS ONE policies on sharing data and materials, as detailed online in the guide for authors.

Author Contributions

Conceived and designed the experiments: GZ FM HS. Performed the experiments: GZ JD P-YS. Analyzed the data: GZ FM JD HS P-YS. Contributed reagents/materials/analysis tools: HS FM JD P-YS RDI. Wrote the paper: GZ HS MB FM.