About the Authors

Cuiling Xu

Contributed equally to this work with: Cuiling Xu, Tian Bai

Affiliation State Key Laboratory for Molecular Virology and Genetic Engineering, National Institute for Viral Infectious Disease Control and Prevention, Chinese Center for Disease Control and Prevention, Beijing, China

Tian Bai

Contributed equally to this work with: Cuiling Xu, Tian Bai

Affiliation State Key Laboratory for Molecular Virology and Genetic Engineering, National Institute for Viral Infectious Disease Control and Prevention, Chinese Center for Disease Control and Prevention, Beijing, China

A. Danielle Iuliano

Affiliation Influenza Division, National Center for Immunization and Respiratory Diseases, Centers for Disease Control and Prevention, Atlanta, Georgia, United States of America

Min Wang

Affiliation State Key Laboratory for Molecular Virology and Genetic Engineering, National Institute for Viral Infectious Disease Control and Prevention, Chinese Center for Disease Control and Prevention, Beijing, China

Lei Yang

Affiliation State Key Laboratory for Molecular Virology and Genetic Engineering, National Institute for Viral Infectious Disease Control and Prevention, Chinese Center for Disease Control and Prevention, Beijing, China

Leying Wen

Affiliation State Key Laboratory for Molecular Virology and Genetic Engineering, National Institute for Viral Infectious Disease Control and Prevention, Chinese Center for Disease Control and Prevention, Beijing, China

Yuhong Zeng

Affiliation State Key Laboratory for Molecular Virology and Genetic Engineering, National Institute for Viral Infectious Disease Control and Prevention, Chinese Center for Disease Control and Prevention, Beijing, China

Xiaodan Li

Affiliation State Key Laboratory for Molecular Virology and Genetic Engineering, National Institute for Viral Infectious Disease Control and Prevention, Chinese Center for Disease Control and Prevention, Beijing, China

Tao Chen

Affiliation State Key Laboratory for Molecular Virology and Genetic Engineering, National Institute for Viral Infectious Disease Control and Prevention, Chinese Center for Disease Control and Prevention, Beijing, China

Wei Wang

Affiliation State Key Laboratory for Molecular Virology and Genetic Engineering, National Institute for Viral Infectious Disease Control and Prevention, Chinese Center for Disease Control and Prevention, Beijing, China

Ying Hu

Affiliation State Key Laboratory for Molecular Virology and Genetic Engineering, National Institute for Viral Infectious Disease Control and Prevention, Chinese Center for Disease Control and Prevention, Beijing, China

Limei Yang

Affiliation State Key Laboratory for Molecular Virology and Genetic Engineering, National Institute for Viral Infectious Disease Control and Prevention, Chinese Center for Disease Control and Prevention, Beijing, China

Zi Li

Affiliation State Key Laboratory for Molecular Virology and Genetic Engineering, National Institute for Viral Infectious Disease Control and Prevention, Chinese Center for Disease Control and Prevention, Beijing, China

Shumei Zou

Affiliation State Key Laboratory for Molecular Virology and Genetic Engineering, National Institute for Viral Infectious Disease Control and Prevention, Chinese Center for Disease Control and Prevention, Beijing, China

Dexin Li

Affiliation State Key Laboratory for Molecular Virology and Genetic Engineering, National Institute for Viral Infectious Disease Control and Prevention, Chinese Center for Disease Control and Prevention, Beijing, China

Shiwen Wang

Affiliation State Key Laboratory for Molecular Virology and Genetic Engineering, National Institute for Viral Infectious Disease Control and Prevention, Chinese Center for Disease Control and Prevention, Beijing, China

Zijian Feng

Affiliation Chinese Center for Disease Control and Prevention, Beijing, China

Yanping Zhang

Affiliation State Key Laboratory for Molecular Virology and Genetic Engineering, National Institute for Viral Infectious Disease Control and Prevention, Chinese Center for Disease Control and Prevention, Beijing, China

Hongjie Yu

Affiliation Chinese Center for Disease Control and Prevention, Beijing, China

Weizhong Yang

Affiliation Chinese Center for Disease Control and Prevention, Beijing, China

Yu Wang

Affiliation Chinese Center for Disease Control and Prevention, Beijing, China

Marc-Alain Widdowson

Affiliation Influenza Division, National Center for Immunization and Respiratory Diseases, Centers for Disease Control and Prevention, Atlanta, Georgia, United States of America

Yuelong Shu

yshu@cnic.org.cn

Affiliation State Key Laboratory for Molecular Virology and Genetic Engineering, National Institute for Viral Infectious Disease Control and Prevention, Chinese Center for Disease Control and Prevention, Beijing, China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: DL SW ZF Y. Zhang WY YW HY YS. Performed the experiments: TB MW LW Y. Zeng XL WW YH ZL SZ. Analyzed the data: CX. Contributed reagents/materials/analysis tools: CX. Wrote the paper: CX TB ADI. Full access to data and responsibility for the integrity of the data and the accuracy of the data analysis: YS. Critical revision of the manuscript for important intellectual content: CX ADI M-AW YS. Administrative, technical, or material support: TB MW Lei Yang LW Y. Zeng XL WW YH LZ SZ TC Limei Yang. Study supervision: SW ZF Y. Zhang.