About the Authors

Takefumi Kikusui

Affiliation Companion Animal Research, School of Veterinary Medicine, Azabu University, Sagamihara, Kanagawa, Japan

Kaori Nakanishi

Affiliation Companion Animal Research, School of Veterinary Medicine, Azabu University, Sagamihara, Kanagawa, Japan

Ryoko Nakagawa

Affiliation Laboratory for Biolinguistics, Mind and Intelligence Research Core, Brain Science Institute (BSI), RIKEN, Wako, Saitama, Japan

Miho Nagasawa

Affiliation Companion Animal Research, School of Veterinary Medicine, Azabu University, Sagamihara, Kanagawa, Japan

Kazutaka Mogi

Affiliation Companion Animal Research, School of Veterinary Medicine, Azabu University, Sagamihara, Kanagawa, Japan

Kazuo Okanoya

okanoya@brain.riken.jp

Affiliations Laboratory for Biolinguistics, Mind and Intelligence Research Core, Brain Science Institute (BSI), RIKEN, Wako, Saitama, Japan, ERATO, Okanoya Emotional Information Project, Japan Science and Technology Corporation, Wako, Saitama, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: TK KN. Performed the experiments: KN RN. Analyzed the data: KN RN MN KM. Wrote the paper: TK KN RN KO.