About the Authors

Eriko Koshimizu

Affiliations Department of Ophthalmology, Harvard Medical School, Boston, Massachusetts, United States of America, Graduate School of Marine Science and Technology, University of Marine Science and Technology, Tokyo, Japan

Shintaro Imamura

Current address: National Research Institute of Fisheries Science, Yokohama, Japan

Affiliation Department of Ophthalmology, Harvard Medical School, Boston, Massachusetts, United States of America

Jie Qi

Current address: Ocean University of China, Qingdao, China

Affiliations Department of Ophthalmology, Harvard Medical School, Boston, Massachusetts, United States of America, Department of Metabolism and Aging, The Scripps Research Institute, Jupiter, Florida, United States of America

Jamal Toure

Affiliation Department of Ophthalmology, Harvard Medical School, Boston, Massachusetts, United States of America

Delgado M. Valdez Jr

Affiliation Department of Metabolism and Aging, The Scripps Research Institute, Jupiter, Florida, United States of America

Christopher E. Carr

Affiliation Department of Earth, Atmospheric and Planetary Sciences, Massachusetts Institute of Technology, Cambridge, Massachusetts, United States of America

Jun-ichi Hanai

Affiliation Division of Nephrology, Interdisciplinary Medicine and Biotechnology, Department of Medicine, Beth Israel Deaconess Medical Center, Harvard Medical School, Boston, Massachusetts, United States of America

Shuji Kishi

kishi@scripps.edu

Affiliations Department of Ophthalmology, Harvard Medical School, Boston, Massachusetts, United States of America, Department of Metabolism and Aging, The Scripps Research Institute, Jupiter, Florida, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: EK SI JH SK. Performed the experiments: EK SI JQ JT DMV SK. Analyzed the data: EK SI CEC SK. Contributed reagents/materials/analysis tools: CEC SK. Wrote the paper: EK SK.