About the Authors

Ikuko Yao

yaoik@takii.kmu.ac.jp (IY); setou@hama-med.ac.jp (MS)

Affiliations Department of Medical Chemistry, Kansai Medical University, Moriguchi, Osaka, Japan, Mitsubishi Kagaku Institute of Life Sciences, Machida, Tokyo, Japan

Keizo Takao

Affiliations Genetic Engineering and Functional Genomics Group, Frontier Technology Center, Graduate School of Medicine Kyoto University, Kyoto, Japan, Section of Behavior Analysis, Center for Genetic Analysis of Behavior, National Institute for Physiological Sciences, Okazaki, Aichi, Japan

Tsuyoshi Miyakawa

Affiliations Genetic Engineering and Functional Genomics Group, Frontier Technology Center, Graduate School of Medicine Kyoto University, Kyoto, Japan, Section of Behavior Analysis, Center for Genetic Analysis of Behavior, National Institute for Physiological Sciences, Okazaki, Aichi, Japan, Division of Systems Medical Science, Institute for Comprehensive Medical Science, Fujita Health University, Toyoake, Aichi, Japan

Seiji Ito

Affiliation Department of Medical Chemistry, Kansai Medical University, Moriguchi, Osaka, Japan

Mitsutoshi Setou

yaoik@takii.kmu.ac.jp (IY); setou@hama-med.ac.jp (MS)

Affiliations Mitsubishi Kagaku Institute of Life Sciences, Machida, Tokyo, Japan, Department of Molecular Anatomy, Hamamatsu University School of Medicine, Hamamatsu, Shizuoka, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: IY TM MS. Performed the experiments: KT IY. Analyzed the data: KT. Contributed reagents/materials/analysis tools: IY TM. Wrote the paper: IY SI MS. Conceived and designed the test battery experiments: TM.