About the Authors

Elizabeth K. Marsh

Affiliation School of Biosciences, University of Birmingham, Birmingham, West Midlands, United Kingdom

Maaike C. W. van den Berg

Affiliation Laboratory of Physiological Chemistry, University Medical Center Utrecht, Utrecht, The Netherlands

Robin C. May

R.C.May@bham.ac.uk

Affiliation School of Biosciences, University of Birmingham, Birmingham, West Midlands, United Kingdom

Competing Interests

RCM is an associate editor for both PLoS One and PLoS Pathogens.

Author Contributions

Conceived and designed the experiments: EKM RCM. Performed the experiments: EKM MCWVDB. Analyzed the data: EKM RCM. Wrote the paper: EKM RCM.