About the Authors

S. Helene Richter

Affiliations Animal Models in Psychiatry, Central Institute of Mental Health (CIMH), Mannheim, Germany, Animal Welfare and Ethology, University of Giessen, Giessen, Germany, Behavioural Biology, University of Muenster, Muenster, Germany

Joseph P. Garner

Affiliation Animal Sciences, Purdue University, West Lafayette, Indiana, United States of America

Benjamin Zipser

Affiliation Behavioural Biology, University of Muenster, Muenster, Germany

Lars Lewejohann

Affiliation Behavioural Biology, University of Muenster, Muenster, Germany

Norbert Sachser

Affiliation Behavioural Biology, University of Muenster, Muenster, Germany

Chadi Touma

Affiliation Psychoneuroendocrinology, Max Planck Institute of Psychiatry, Munich, Germany

Britta Schindler

Affiliation Psychoneuroendocrinology, Max Planck Institute of Psychiatry, Munich, Germany

Sabine Chourbaji

Affiliation Animal Models in Psychiatry, Central Institute of Mental Health (CIMH), Mannheim, Germany

Christiane Brandwein

Affiliation Animal Models in Psychiatry, Central Institute of Mental Health (CIMH), Mannheim, Germany

Peter Gass

Affiliation Animal Models in Psychiatry, Central Institute of Mental Health (CIMH), Mannheim, Germany

Niek van Stipdonk

Affiliation Delta Phenomics BV, Utrecht, The Netherlands

Johanneke van der Harst

Affiliation Delta Phenomics BV, Utrecht, The Netherlands

Berry Spruijt

Affiliation Delta Phenomics BV, Utrecht, The Netherlands

Vootele Võikar

Affiliations Neuroscience Center, University of Helsinki, Helsinki, Finland, Institute of Anatomy, University of Zürich, Zürich, Switzerland

David P. Wolfer

Affiliation Institute of Anatomy, University of Zürich, Zürich, Switzerland

Hanno Würbel

hanno.wuerbel@vetmed.uni-giessen.de

Affiliation Animal Welfare and Ethology, University of Giessen, Giessen, Germany

Competing Interests

Three authors (Niek van Stipdonk, Johanneke van der Harst and Berry Spruijt) are employed by a commercial company (Delta Phenomics BV), and part of the study was conducted in a lab of this company. However, the company played no other role in this study. The collaboration was based on Berry Spruijt's expertise in behavioural phenotyping and not on anything related to the company. Moreover, the costs for the study (animals, consumables) and the costs for the biotechnician who tested the animals was covered by the grant provided by the German Research Foundation (DFG). The authors confirm that the affiliation to this company does not alter their adherence to all the PLoS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: SHR HW. Performed the experiments: SHR BZ CT BS SC CB NvS VV. Analyzed the data: SHR HW JPG. Contributed reagents/materials/analysis tools: HW DPW PG BS NS CT. Wrote the paper: SHR HW JPG. Supervised the experimental work in the Muenster lab: LL. Supervised the experimental work in the Utrecht lab: JvdH. Contributed to manuscript preparation: LL JvdH B. Spruijt. Provided the lab space and equipment in the Utrecht lab, organised the experimental part of this lab: B. Spruijt. Conducted part of the behavioural testing in the Munich lab: B. Schindler.