About the Authors

Heather D. VanGuilder

Contributed equally to this work with: Heather D. VanGuilder, Georgina V. Bixler

Affiliation Department of Pharmacology, Penn State College of Medicine, Hershey, Pennsylvania, United States of America

Georgina V. Bixler

Contributed equally to this work with: Heather D. VanGuilder, Georgina V. Bixler

Affiliation Department of Pharmacology, Penn State College of Medicine, Hershey, Pennsylvania, United States of America

Lydia Kutzler

Affiliation Department of Cellular & Molecular Physiology, Penn State College of Medicine, Hershey, Pennsylvania, United States of America

Robert M. Brucklacher

Affiliation Department of Pharmacology, Penn State College of Medicine, Hershey, Pennsylvania, United States of America

Sarah K. Bronson

Affiliation Department of Cellular & Molecular Physiology, Penn State College of Medicine, Hershey, Pennsylvania, United States of America

Scot R. Kimball

Affiliation Department of Cellular & Molecular Physiology, Penn State College of Medicine, Hershey, Pennsylvania, United States of America

Willard M. Freeman

wfreeman@psu.edu

Affiliation Department of Pharmacology, Penn State College of Medicine, Hershey, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: WMF SRK SKB HDV. Performed the experiments: HDV GVB LK RMB. Analyzed the data: HDV GVB SKB WMF. Wrote the paper: HDV GVB RMB SKB SRK WMF.