About the Authors

Daisuke Matsumaru

Affiliation Global COE "Cell Fate Regulation Research and Education Unit", Department of Organ Formation, Institute of Molecular Embryology and Genetics (IMEG), Kumamoto University, Kumamoto, Japan

Ryuma Haraguchi

Current address: Department of Molecular Pathology, Ehime University Graduate School of Medicine, Ehime, Japan

Affiliation Global COE "Cell Fate Regulation Research and Education Unit", Department of Organ Formation, Institute of Molecular Embryology and Genetics (IMEG), Kumamoto University, Kumamoto, Japan

Shinichi Miyagawa

Current address: Okazaki Institute for Integrative Bioscience, National Institutes of Natural Sciences, Aichi, Japan

Affiliation Global COE "Cell Fate Regulation Research and Education Unit", Department of Organ Formation, Institute of Molecular Embryology and Genetics (IMEG), Kumamoto University, Kumamoto, Japan

Jun Motoyama

Affiliation Department of Medical Life Systems, Doshisha University, Kyoto, Japan

Naomi Nakagata

Affiliation Center for Animal Resources and Development (CARD), Kumamoto University, Kumamoto, Japan

Frits Meijlink

Affiliation Hubrecht Institute, KNAW and University Medical Center, Utrecht, The Netherlands

Gen Yamada

gensan@gpo.kumamoto-u.ac.jp

Affiliation Global COE "Cell Fate Regulation Research and Education Unit", Department of Organ Formation, Institute of Molecular Embryology and Genetics (IMEG), Kumamoto University, Kumamoto, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: DM RH SM GY. Performed the experiments: DM RH SM JM NN. Analyzed the data: DM RH FM GY. Wrote the paper: DM GY.