About the Authors

Sean M. McMahon

mcmahons@si.edu

Affiliation Smithsonian Tropical Research Institute, Center for Tropical Forest Science, Smithsonian Environmental Research Center, Edgewater, Maryland, United States of America

Charlotte J. E. Metcalf

Affiliation Department of Zoology, Oxford University, Oxford, United Kingdom

Christopher W. Woodall

Affiliation Forest Inventory and Analysis, U.S. Forest Service, St. Paul, Minnesota, United States of America

Competing Interests

SMM was funded by HSBC Bank as a postdoctoral fellow through the Smithsonian Institution. This does not alter the authors' adherence to all the PLoS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: SMM CJM. Analyzed the data: SMM CJM. Contributed reagents/materials/analysis tools: CW. Wrote the paper: SMM CJM CW.