About the Authors

Paul Plummer

pplummer@iastate.edu

Affiliations Department of Veterinary Microbiology and Preventative Medicine, College of Veterinary Medicine, Iowa State University, Ames, Iowa, United States of America, Department of Veterinary Diagnostics and Production Animal Medicine, College of Veterinary Medicine, Iowa State University, Ames, Iowa, United States of America

Jinge Zhu

Affiliation Department of Chemistry, The Ohio State University, Columbus, Ohio, United States of America

Masato Akiba

Current address: National Institute of Animal Health, Tsukuba, Ibaraki, Japan

Affiliation Department of Veterinary Microbiology and Preventative Medicine, College of Veterinary Medicine, Iowa State University, Ames, Iowa, United States of America

Dehua Pei

Affiliation Department of Chemistry, The Ohio State University, Columbus, Ohio, United States of America

Qijing Zhang

Affiliation Department of Veterinary Microbiology and Preventative Medicine, College of Veterinary Medicine, Iowa State University, Ames, Iowa, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: PP MA JZ DP QZ. Performed the experiments: PP MA JZ. Analyzed the data: PP MA JZ DP QJ. Contributed reagents/materials/analysis tools: PP JZ MA DP QZ. Wrote the paper: PP DP QZ.