About the Authors

David J. Wardle

Contributed equally to this work with: David J. Wardle, Joseph Burgon

Affiliation Medical Research Council Centre for Developmental and Biomedical Genetics, University of Sheffield, Sheffield, United Kingdom

Joseph Burgon

Contributed equally to this work with: David J. Wardle, Joseph Burgon

Affiliation Medical Research Council Centre for Developmental and Biomedical Genetics, University of Sheffield, Sheffield, United Kingdom

Ian Sabroe

Affiliation Department of Infection and Immunity, University of Sheffield, Sheffield, United Kingdom

Colin D. Bingle

Affiliation Department of Infection and Immunity, University of Sheffield, Sheffield, United Kingdom

Moira K. B. Whyte

Affiliations Medical Research Council Centre for Developmental and Biomedical Genetics, University of Sheffield, Sheffield, United Kingdom, Department of Infection and Immunity, University of Sheffield, Sheffield, United Kingdom

Stephen A. Renshaw

s.a.renshaw@sheffield.ac.uk

Affiliations Medical Research Council Centre for Developmental and Biomedical Genetics, University of Sheffield, Sheffield, United Kingdom, Department of Infection and Immunity, University of Sheffield, Sheffield, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: IS CDB MKBW SAR. Performed the experiments: DJW JB. Analyzed the data: SW JB SAR. Wrote the paper: DJW IS CDB MKBW SAR.