About the Authors

Hsiao-Chun Huang

Affiliations Department of Systems Biology, Harvard Medical School, Boston, Massachusetts, United States of America, Graduate Program in Systems Biology, Harvard Medical School, Boston, Massachusetts, United States of America

Timothy J. Mitchison

Affiliation Department of Systems Biology, Harvard Medical School, Boston, Massachusetts, United States of America

Jue Shi

jshi@hkbu.eduk.hk

Affiliations Department of Systems Biology, Harvard Medical School, Boston, Massachusetts, United States of America, Center for Quantitative Systems Biology and Department of Physics, Hong Kong Baptist University, Kowloon Tong, Hong Kong

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: H-CH TJM JS. Performed the experiments: H-CH. Analyzed the data: H-CH JS. Contributed reagents/materials/analysis tools: H-CH TJM JS. Wrote the paper: H-CH TJM JS.