About the Authors

Girishchandra B. Patel

girish.patel@nrc-cnrc.gc.ca

Affiliation Institute for Biological Sciences, National Research Council Canada, Ottawa, Canada

Hongyan Zhou

Affiliation Institute for Biological Sciences, National Research Council Canada, Ottawa, Canada

Amalia Ponce

Affiliation Institute for Biological Sciences, National Research Council Canada, Ottawa, Canada

Greg Harris

Affiliation Institute for Biological Sciences, National Research Council Canada, Ottawa, Canada

Wangxue Chen

Affiliation Institute for Biological Sciences, National Research Council Canada, Ottawa, Canada

Competing Interests

GBP and WC are co-inventors on patent applications (owned by the Crown/Government of Canada) on the archaeal lipid mucosal vaccine adjuvant and delivery (AMVAD) technology. None of the other authors declare any conflicts. This does not alter the authors' adherence to all the PLoS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: GBP WC. Performed the experiments: HZ AP GH. Analyzed the data: GBP HZ AP GH WC. Wrote the paper: GBP WC.