About the Authors

Lauren Lederle

Contributed equally to this work with: Lauren Lederle, Susanna Weber, Tara Wright

Affiliation Section on Behavioral Science and Genetics, Laboratory for Integrative Neuroscience, National Institute on Alcohol Abuse and Alcoholism, National Institutes of Health, Rockville, Maryland, United States of America

Susanna Weber

Contributed equally to this work with: Lauren Lederle, Susanna Weber, Tara Wright

Affiliation Section on Behavioral Science and Genetics, Laboratory for Integrative Neuroscience, National Institute on Alcohol Abuse and Alcoholism, National Institutes of Health, Rockville, Maryland, United States of America

Tara Wright

Contributed equally to this work with: Lauren Lederle, Susanna Weber, Tara Wright

Affiliation Section on Behavioral Science and Genetics, Laboratory for Integrative Neuroscience, National Institute on Alcohol Abuse and Alcoholism, National Institutes of Health, Rockville, Maryland, United States of America

Michael Feyder

Affiliation Section on Behavioral Science and Genetics, Laboratory for Integrative Neuroscience, National Institute on Alcohol Abuse and Alcoholism, National Institutes of Health, Rockville, Maryland, United States of America

Jonathan L. Brigman

Affiliation Section on Behavioral Science and Genetics, Laboratory for Integrative Neuroscience, National Institute on Alcohol Abuse and Alcoholism, National Institutes of Health, Rockville, Maryland, United States of America

Hans S. Crombag

Affiliation Behavioural and Clinical Neuroscience Research Group, School of Psychology, University of Sussex, Brighton, United Kingdom

Lisa M. Saksida

Affiliation Department of Experimental Psychology and the Medical Research Council and Wellcome Trust Behavioral and Clinical Neuroscience Institute, University of Cambridge, Cambridge, United Kingdom

Timothy J. Bussey

Affiliation Department of Experimental Psychology and the Medical Research Council and Wellcome Trust Behavioral and Clinical Neuroscience Institute, University of Cambridge, Cambridge, United Kingdom

Andrew Holmes

holmesan@mail.nih.gov

Affiliation Section on Behavioral Science and Genetics, Laboratory for Integrative Neuroscience, National Institute on Alcohol Abuse and Alcoholism, National Institutes of Health, Rockville, Maryland, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: AH LL SW MF TW. Performed the experiments: LL SW MF TW. Analyzed the data: LL AH SW MF TW. Contributed reagents/materials/analysis tools: TJB LMS HSC. Wrote the paper: LL SW TW MF TJB HSC AH JLB.