About the Authors

Lumine Matsumoto

Affiliation Division of Neuroscience, Department of Neurology, Graduate School of Medicine, The University of Tokyo, Bunkyo, Tokyo, Japan

Hiroshi Takuma

Affiliation Department of Neurology, University of Tsukuba, Tsukuba, Ibaraki, Japan

Akira Tamaoka

Affiliation Department of Neurology, University of Tsukuba, Tsukuba, Ibaraki, Japan

Hiroshi Kurisaki

Affiliation Department of Neurology, National Hospital Organization Tokyo Hospital, Kiyose, Tokyo, Japan

Hidetoshi Date

Affiliation Division of Neuroscience, Department of Neurology, Graduate School of Medicine, The University of Tokyo, Bunkyo, Tokyo, Japan

Shoji Tsuji

Affiliation Division of Neuroscience, Department of Neurology, Graduate School of Medicine, The University of Tokyo, Bunkyo, Tokyo, Japan

Atsushi Iwata

iwata-tky@umin.ac.jp

Affiliations Division of Neuroscience, Department of Neurology, Graduate School of Medicine, The University of Tokyo, Bunkyo, Tokyo, Japan, Department of Molecular Neuroscience on Neurodegeneration, Graduate School of Medicine, The University of Tokyo, Bunkyo, Tokyo, Japan, PRESTO, Japan Science and Technology Agency (JST), Kawaguchi, Saitama, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: LM ST AI. Performed the experiments: LM HD AI. Analyzed the data: LM AI. Contributed reagents/materials/analysis tools: HT AT HK ST. Wrote the paper: LM HT AI.