About the Authors

Kristen B. Greenland

Affiliations Department of Biochemistry, University of Washington, Seattle, Washington, United States of America, Program in Molecular and Cellular Biology, University of Washington, Seattle, Washington, United States of America

Huiming Ding

Affiliation Banting and Best Department of Medical Research, Terrence Donnelly Centre for Cellular and Biomolecular Research, University of Toronto, Toronto, Ontario, Canada

Michael Costanzo

Affiliation Banting and Best Department of Medical Research, Terrence Donnelly Centre for Cellular and Biomolecular Research, University of Toronto, Toronto, Ontario, Canada

Charles Boone

Affiliation Banting and Best Department of Medical Research, Terrence Donnelly Centre for Cellular and Biomolecular Research, University of Toronto, Toronto, Ontario, Canada

Trisha N. Davis

tdavis@u.washington.edu

Affiliations Department of Biochemistry, University of Washington, Seattle, Washington, United States of America, Program in Molecular and Cellular Biology, University of Washington, Seattle, Washington, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: KG TD CB MC. Performed the experiments: KG. Analyzed the data: KG TD MC HD. Contributed reagents/materials/analysis tools: HD. Wrote the paper: KG TD.