About the Authors

Mairi H. McLean

Affiliation Gastrointestinal Research Group, School of Medicine and Dentistry, Aberdeen University, Aberdeen, United Kingdom

Graeme I. Murray

Affiliation Department of Pathology, Aberdeen University, Aberdeen, United Kingdom

Keith N. Stewart

Affiliation Department of Pathology, Aberdeen University, Aberdeen, United Kingdom

Gillian Norrie

Affiliation Colorectal Surgery Unit, Western General Hospital, Edinburgh, United Kingdom

Claus Mayer

Affiliation Biomathematics and Statistics Scotland, The Rowett Institute of Nutrition and Health, Aberdeen University, Aberdeen, United Kingdom

Georgina L. Hold

Affiliation Gastrointestinal Research Group, School of Medicine and Dentistry, Aberdeen University, Aberdeen, United Kingdom

John Thomson

Affiliation Gastrointestinal Research Group, School of Medicine and Dentistry, Aberdeen University, Aberdeen, United Kingdom

Nicky Fyfe

Affiliation Department of Pathology, Aberdeen University, Aberdeen, United Kingdom

Mairi Hope

Affiliation Gastrointestinal Research Group, School of Medicine and Dentistry, Aberdeen University, Aberdeen, United Kingdom

N. Ashley G. Mowat

Affiliation Gastrointestinal Research Group, School of Medicine and Dentistry, Aberdeen University, Aberdeen, United Kingdom

Janice E. Drew

Affiliation The Rowett Institute of Nutrition and Health, Aberdeen University, Aberdeen, United Kingdom

Emad M. El-Omar

e.el-omar@abdn.ac.uk

Affiliation Gastrointestinal Research Group, School of Medicine and Dentistry, Aberdeen University, Aberdeen, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: MHM EEO GIM. Performed the experiments: MHM KNS JED NF GLH JT MH. Analyzed the data: GN CM MHM EEO GIM JED JT NAGM. Contributed reagents/materials/analysis tools: MHM GIM KNS NF JT JED. Wrote the paper: MHM EEO GIM.