About the Authors

Yoon Ha Choi

Affiliation Division of Molecular and Life Science, Department of Life Science, Pohang University of Science and Technology (POSTECH), Pohang, Republic of Korea

Choon-Ho Park

Affiliation Division of Molecular and Life Science, Department of Life Science, Pohang University of Science and Technology (POSTECH), Pohang, Republic of Korea

Wanil Kim

Affiliation Division of Molecular and Life Science, Department of Life Science, Pohang University of Science and Technology (POSTECH), Pohang, Republic of Korea

Hua Ling

Affiliation School of Chemical and Biomedical Engineering, Nanyang Technological University, Singapore, Singapore

Aram Kang

Affiliation School of Chemical and Biomedical Engineering, Nanyang Technological University, Singapore, Singapore

Matthew Wook Chang

Affiliation School of Chemical and Biomedical Engineering, Nanyang Technological University, Singapore, Singapore

Sun-Kyoung Im

Affiliation Division of Molecular and Life Science, Department of Life Science, Pohang University of Science and Technology (POSTECH), Pohang, Republic of Korea

Hyun-Woo Jeong

Affiliation Division of Molecular and Life Science, Department of Life Science, Pohang University of Science and Technology (POSTECH), Pohang, Republic of Korea

Young-Yun Kong

Affiliation Department of Biological Sciences, Seoul National University, Seoul, Republic of Korea

Kyong-Tai Kim

ktk@postech.ac.kr

Affiliations Division of Molecular and Life Science, Department of Life Science, Pohang University of Science and Technology (POSTECH), Pohang, Republic of Korea, Division of Integrative Bioscience and Biotechnology, Pohang University of Science and Technology (POSTECH), Pohang, Republic of Korea

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: YHC MWC SKI KTK. Performed the experiments: YHC CHP WK HL AK. Analyzed the data: YHC HWJ. Contributed reagents/materials/analysis tools: KTK. Wrote the paper: YHC KTK. Made VRK1 knock-out mice and gave valuable discussion: YYK.