About the Authors

Xinran Xiang

Contributed equally to this work with: Xinran Xiang, Yen Phung

Affiliation Laboratory of Molecular Biology, Center for Cancer Research, National Cancer Institute, National Institutes of Health, Bethesda, Maryland, United States of America

Yen Phung

Contributed equally to this work with: Xinran Xiang, Yen Phung

Affiliation Laboratory of Molecular Biology, Center for Cancer Research, National Cancer Institute, National Institutes of Health, Bethesda, Maryland, United States of America

Mingqian Feng

Affiliation Laboratory of Molecular Biology, Center for Cancer Research, National Cancer Institute, National Institutes of Health, Bethesda, Maryland, United States of America

Kunio Nagashima

Affiliation Science Applications International Corporation-Frederick, Inc., Frederick, Maryland, United States of America

Jingli Zhang

Affiliation Laboratory of Molecular Biology, Center for Cancer Research, National Cancer Institute, National Institutes of Health, Bethesda, Maryland, United States of America

V. Courtney Broaddus

Affiliation Lung Biology Center, University of California San Francisco, San Francisco, California, United States of America

Raffit Hassan

Affiliation Laboratory of Molecular Biology, Center for Cancer Research, National Cancer Institute, National Institutes of Health, Bethesda, Maryland, United States of America

David FitzGerald

Affiliation Laboratory of Molecular Biology, Center for Cancer Research, National Cancer Institute, National Institutes of Health, Bethesda, Maryland, United States of America

Mitchell Ho

homi@mail.nih.gov

Affiliation Laboratory of Molecular Biology, Center for Cancer Research, National Cancer Institute, National Institutes of Health, Bethesda, Maryland, United States of America

Competing Interests

KN is employed at SAIC-Frederick, but is funded by NCI. SAIC provided no funds for this research. KN provided analytical support. This does not alter the authors' adherence to all the PLoS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: VCB DF MH. Performed the experiments: XX YP KN. Analyzed the data: XX YP KN DF MH. Contributed reagents/materials/analysis tools: MF JZ RH. Wrote the paper: XX YP MH. Scientific supervision: MH.