About the Authors

Jae-Min Song

Affiliation Department of Microbiology and Immunology and Emory Vaccine Center, Emory University School of Medicine, Atlanta, Georgia, United States of America

Bao-Zhong Wang

Affiliation Department of Microbiology and Immunology and Emory Vaccine Center, Emory University School of Medicine, Atlanta, Georgia, United States of America

Kyoung-Mi Park

Affiliation Department of Microbiology and Immunology and Emory Vaccine Center, Emory University School of Medicine, Atlanta, Georgia, United States of America

Nico Van Rooijen

Affiliation Department of Molecular Cell Biology, Vrije Universiteit Medisch Centrum, Amsterdam, The Netherlands

Fu-Shi Quan

Affiliation Department of Microbiology and Immunology and Emory Vaccine Center, Emory University School of Medicine, Atlanta, Georgia, United States of America

Min-Chul Kim

Affiliation Department of Microbiology and Immunology and Emory Vaccine Center, Emory University School of Medicine, Atlanta, Georgia, United States of America

Hyun-Tak Jin

Affiliation Department of Microbiology and Immunology and Emory Vaccine Center, Emory University School of Medicine, Atlanta, Georgia, United States of America

Andrew Pekosz

Affiliation Department of Molecular Microbiology and Immunology, Johns Hopkins University, Baltimore, Maryland, United States of America

Richard W. Compans

rcompan@emory.edu (RWC); skang2@emory.edu (SMK)

Affiliation Department of Microbiology and Immunology and Emory Vaccine Center, Emory University School of Medicine, Atlanta, Georgia, United States of America

Sang-Moo Kang

rcompan@emory.edu (RWC); skang2@emory.edu (SMK)

Affiliation Department of Microbiology and Immunology and Emory Vaccine Center, Emory University School of Medicine, Atlanta, Georgia, United States of America

Competing Interests

S.M.K., R.W.C. and Emory University have equity interests in Zetra Biologicals, which is developing virus-like particle technology under license from Emory University. This does not alter the authors' adherence to all the PLoS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: JMS RWC SMK. Performed the experiments: JMS KMP MCK. Analyzed the data: JMS SMK. Contributed reagents/materials/analysis tools: BZW NVR FSQ HTJ AP SMK. Wrote the paper: JMS RWC SMK.