About the Authors

Fumio Yoshikawa

Affiliation Laboratory for Molecular Neurogenesis, RIKEN Brain Science Institute, Wako, Saitama, Japan

Yoshiko Banno

Affiliation Department of Cell Signaling, Gifu University Graduate School of Medicine, Gifu, Gifu, Japan

Yoshinori Otani

Affiliation Department of Molecular Neurobiology, Tokyo University of Pharmacy and Life Sciences, Hachioji, Tokyo, Japan

Yoshihide Yamaguchi

Affiliation Department of Molecular Neurobiology, Tokyo University of Pharmacy and Life Sciences, Hachioji, Tokyo, Japan

Yuko Nagakura-Takagi

Affiliation Laboratory for Molecular Neurogenesis, RIKEN Brain Science Institute, Wako, Saitama, Japan

Noriyuki Morita

Affiliations Laboratory for Molecular Neurogenesis, RIKEN Brain Science Institute, Wako, Saitama, Japan, Yasuda Women's University, Hiroshima, Hiroshima, Japan

Yumi Sato

Affiliation Laboratory for Molecular Neurogenesis, RIKEN Brain Science Institute, Wako, Saitama, Japan

Chihiro Saruta

Affiliation Laboratory for Molecular Neurogenesis, RIKEN Brain Science Institute, Wako, Saitama, Japan

Hirozumi Nishibe

Affiliation Laboratory for Molecular Neurogenesis, RIKEN Brain Science Institute, Wako, Saitama, Japan

Tetsushi Sadakata

Affiliations Laboratory for Molecular Neurogenesis, RIKEN Brain Science Institute, Wako, Saitama, Japan, JST, CREST, Kawaguchi, Saitama, Japan

Yo Shinoda

Current address: European Neuroscience Institute Göttingen, Göttingen, Germany

Affiliations Laboratory for Molecular Neurogenesis, RIKEN Brain Science Institute, Wako, Saitama, Japan, JST, CREST, Kawaguchi, Saitama, Japan

Kanehiro Hayashi

Current address: Research Institute Pharmaceutical Science, Musashino University, Nishitokyo, Tokyo, Japan

Affiliations Laboratory for Molecular Neurogenesis, RIKEN Brain Science Institute, Wako, Saitama, Japan, JST, CREST, Kawaguchi, Saitama, Japan

Yuriko Mishima

Affiliations Laboratory for Molecular Neurogenesis, RIKEN Brain Science Institute, Wako, Saitama, Japan, JST, CREST, Kawaguchi, Saitama, Japan

Hiroko Baba

Affiliation Department of Molecular Neurobiology, Tokyo University of Pharmacy and Life Sciences, Hachioji, Tokyo, Japan

Teiichi Furuichi

tfuruichi@brain.riken.jp

Affiliations Laboratory for Molecular Neurogenesis, RIKEN Brain Science Institute, Wako, Saitama, Japan, JST, CREST, Kawaguchi, Saitama, Japan, Saitama University Graduate School of Science and Engineering, Saitama, Saitama, Japan, Hiroshima University Graduate School of Biomedical Sciences, Hiroshima, Hiroshima, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: FY YB YO YY HB TF. Performed the experiments: FY YB YO YNT NM YS CS HN. Analyzed the data: FY YB YO YY HN TS YS KH YM TF. Wrote the paper: YB YY HB TF.