About the Authors

Kazuki Miyaji

Affiliation Department of Animal Science and Biotechnology, Azabu University, Sagamihara, Japan

Kenji Nagao

Affiliation Institute of Life Sciences, Ajinomoto Co. Inc., Kawasaki, Japan

Makoto Bannai

makoto_bannai@ajinomoto.com

Affiliation Institute of Life Sciences, Ajinomoto Co. Inc., Kawasaki, Japan

Hiroshi Asakawa

Affiliation Department of Animal Care, Shimoda Floating Aquarium, Shimoda, Japan

Kaoru Kohyama

Affiliation Department of Animal Care and Management, Izu-Mito Sea Paradise, Numazu, Japan

Dai Ohtsu

Affiliation Aqua Resorts, Yokohama-Hakkeijima Sea Paradise, Yokohama, Japan

Fumio Terasawa

Affiliation Enoshima Aquarium, Fujisawa, Japan

Shu Ito

Affiliation Veterinary Hospital, Adventure World, Nishimuro, Japan

Hajime Iwao

Affiliation Exhibition Division, Niigata City Aquarium Marinepia Nihonkai, Niigata, Japan

Nobuyo Ohtani

Affiliation Department of Animal Science and Biotechnology, Azabu University, Sagamihara, Japan

Mitsuaki Ohta

Affiliation Department of Animal Science and Biotechnology, Azabu University, Sagamihara, Japan

Competing Interests

Kenji Nagao and Makoto Bannai are employees of Ajinomoto Co., Inc. Hiroshi Asakawa is an employee of Shimoda Floating Aquarium. Kaoru Kohyama is an employee of Izu-Mito Sea Paradise. Dai Ohtsu is an employee of Yokohama-Hakkeijima Sea Paradise. Fumio Terasawa is an employee of Enoshima Aquarium. Shu Ito is an employee of Adventure World. Hajime Iwao is an employee of Niigata City Aquarium Marinepia Nihonkai. This does not alter the authors' adherence to all the PLoS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: MB NO MO. Performed the experiments: KM HA KK DO FT SI HI. Analyzed the data: KN MB. Wrote the paper: KM KN MB.