About the Authors

Claudio Ottoni

Contributed equally to this work with: Claudio Ottoni, Giuseppina Primativo

Affiliations Department of Biology, University of Rome Tor Vergata, Rome, Italy, Laboratory of Forensic Genetics and Molecular Archaeology, Universitaire Ziekenhuizen, Leuven, Belgium, Center for Archaeological Sciences, Katholieke Universiteit Leuven, Leuven, Belgium, Center for Human Genetics, Katholieke Universiteit Leuven, Leuven, Belgium

Giuseppina Primativo

Contributed equally to this work with: Claudio Ottoni, Giuseppina Primativo

Affiliation Department of Biology, University of Rome Tor Vergata, Rome, Italy

Baharak Hooshiar Kashani

Affiliation Dipartimento di Genetica e Microbiologia, Università di Pavia, Pavia, Italy

Alessandro Achilli

Affiliations Dipartimento di Genetica e Microbiologia, Università di Pavia, Pavia, Italy, Dipartimento di Biologia Cellulare e Ambientale, Università di Perugia, Perugia, Italy

Cristina Martínez-Labarga

Affiliation Department of Biology, University of Rome Tor Vergata, Rome, Italy

Gianfranco Biondi

Affiliation Dipartimento di Scienze Ambientali, Università dell'Aquila, L'Aquila, Italy

Antonio Torroni

Affiliation Dipartimento di Genetica e Microbiologia, Università di Pavia, Pavia, Italy

Olga Rickards

rickards@uniroma2.it

Affiliation Department of Biology, University of Rome Tor Vergata, Rome, Italy

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: CO AA CML GB AT OR. Performed the experiments: GP BHK AA CML. Analyzed the data: CO GP BHK AA AT. Contributed reagents/materials/analysis tools: GB AT OR. Wrote the paper: CO GP AA AT.