About the Authors

Rebecca Williams

Affiliation Cardiff School of Biosciences, Cardiff University, Cardiff, United Kingdom

Ilyas M. Khan

Affiliation Cardiff School of Biosciences, Cardiff University, Cardiff, United Kingdom

Kirsty Richardson

Affiliation Cardiff School of Biosciences, Cardiff University, Cardiff, United Kingdom

Larissa Nelson

Affiliation Cardiff School of Biosciences, Cardiff University, Cardiff, United Kingdom

Helen E. McCarthy

Affiliation Cardiff School of Biosciences, Cardiff University, Cardiff, United Kingdom

Talal Analbelsi

Affiliation Cardiff School of Biosciences, Cardiff University, Cardiff, United Kingdom

Sim K. Singhrao

Affiliation Cardiff School of Biosciences, Cardiff University, Cardiff, United Kingdom

Gary P. Dowthwaite

Affiliation Cardiff School of Biosciences, Cardiff University, Cardiff, United Kingdom

Rhiannon E. Jones

Affiliation Department of Pathology, Cardiff University, Cardiff, United Kingdom

Duncan M. Baird

Affiliation Department of Pathology, Cardiff University, Cardiff, United Kingdom

Holly Lewis

Affiliation Cytogenetics Department, University Hospital of Wales, Cardiff, United Kingdom

Selwyn Roberts

Affiliation Cytogenetics Department, University Hospital of Wales, Cardiff, United Kingdom

Hannah M. Shaw

Affiliation Cardiff School of Biosciences, Cardiff University, Cardiff, United Kingdom

Jayesh Dudhia

Affiliation Department of Veterinary Clinical Sciences, The Royal Veterinary College, North Mymms, United Kingdom

John Fairclough

Affiliation Department of Orthopaedics, University Hospital of Wales, Cardiff, United Kingdom

Timothy Briggs

Affiliation Royal National Orthopaedic Hospital, Stanmore, United Kingdom

Charles W. Archer

archer@cf.ac.uk

Affiliation Cardiff School of Biosciences, Cardiff University, Cardiff, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: RW IMK HEM CWA. Performed the experiments: RW IMK KR LN HEM TA SKS REJ DMB HL SR GPD. Analyzed the data: RW IMK KR HEM SKS DMB HL SR HMS CWA. Contributed reagents/materials/analysis tools: RW DMB SR JD JF TB CWA. Wrote the paper: RW IMK HEM SKS DMB SR CWA.